- B.Sc. 1981-1984 The Hebrew University of Jerusalem, Faculty of Agriculture, Department of Animal Physiology. M.Sc. 1984-1986 Weizmann Institute of Science, Department of Chemical Immunology, supervised by Prof. Zvi Fishelzon. "Identification of the Regulatory Site of the Complement Protein Factor-H". Ph.D. 1987-1992 Weizmann Institute of Science, Department of Biochemistry, supervised by Prof. Uri Pick. "Structure Function and Regulation of The Plasma Membrane Proton Pump From the Acidophylic Alga- Dunaliella-Acidophila".